Subspecies of Hypolepis rugosula (Dennstaedtiaceae; Pteridophyta) around the world: morphological and biogeographic perspectives

The "Hypolepis rugosula complex" has been the subject of great debate among pteridologists: some have considered H. rugosula a single subcosmopolitan (or circum-Antarctic) species, whereas others have considered it a species-complex, encompassing several species. In the 1920s and 1930s, four geographically distinct varietiesof H. rugosula were recognized. In this work, we present a new taxonomy (with new combinations and statuses, as well as typification and full synonymy), together with complete distribution data for the species, with an infraspecific classification based on morphological and...